GREMLIN Database
GLPD - Aerobic glycerol-3-phosphate dehydrogenase
UniProt: P13035 Sequences:
Download Alignment
We filter this alignment to remove sites that have many gaps before running GREMLIN.

Date: 2015_01
EcoGene: EG10394
Length: 501 (475)
Sequences: 3648 (2403)
Seq/√Len: 110.3

GREMLIN Results:

Legend: The darker and larger the blue dots, the higher coevolution strength.

Residue pairs sorted by coevolution strength:
i j Sco Prob
7_I184_V5.7481.00
4_K30_L4.1291.00
34_A171_R3.9941.00
128_N138_G3.9411.00
32_L186_A3.7761.00
159_V164_G3.6391.00
187_E194_K3.5331.00
236_V268_I3.4811.00
286_E306_R3.3921.00
245_A248_L3.3111.00
290_N294_N3.2751.00
171_R189_I3.2701.00
32_L172_A3.2631.00
186_A197_W3.2081.00
185_E194_K3.1171.00
17_G358_Y3.0981.00
35_Q39_C3.0631.00
170_T186_A3.0101.00
7_I199_A2.9911.00
185_E196_S2.9781.00
4_K165_E2.8911.00
37_L168_T2.8331.00
7_I197_W2.8291.00
190_D461_R2.8211.00
290_N306_R2.8141.00
174_S185_E2.7591.00
232_S292_L2.7231.00
460_V466_A2.7101.00
188_D191_T2.7051.00
16_A156_A2.6461.00
203_V369_L2.6161.00
173_T187_E2.5821.00
19_A29_V2.5331.00
171_R187_E2.4401.00
448_E452_K2.3761.00
176_R183_I2.3661.00
448_E488_W2.3621.00
30_L197_W2.3261.00
11_G31_M2.3041.00
7_I30_L2.2591.00
297_N304_L2.2551.00
427_L457_H2.2491.00
206_T213_F2.2301.00
451_L466_A2.1921.00
227_R274_E2.1031.00
9_I213_F2.0881.00
156_A166_V2.0861.00
13_I155_N2.0851.00
424_N453_Y2.0621.00
420_T471_T2.0551.00
244_Q248_L1.9871.00
286_E290_N1.9821.00
18_I366_L1.9171.00
157_Q424_N1.8721.00
34_A280_K1.8711.00
176_R196_S1.8521.00
440_E448_E1.8471.00
160_V166_V1.8281.00
175_A202_L1.8121.00
478_N481_Q1.8021.00
8_V18_I1.7801.00
451_L485_V1.7791.00
11_G33_E1.7791.00
6_L201_G1.7741.00
16_A152_V1.7681.00
293_L304_L1.7021.00
125_F128_N1.7011.00
5_D200_R1.6941.00
284_I312_T1.6901.00
161_R424_N1.6811.00
201_G369_L1.6801.00
176_R182_W1.6791.00
32_L184_V1.6761.00
24_G380_W1.6581.00
20_A158_M1.6351.00
405_R426_E1.6161.00
338_I372_Y1.6151.00
230_K273_V1.6141.00
289_I312_T1.6121.00
7_I32_L1.6071.00
181_L202_L1.6071.00
21_D381_T1.6071.00
454_L460_V1.5951.00
454_L466_A1.5941.00
289_I306_R1.5931.00
24_G384_S1.5921.00
121_T140_E1.5901.00
18_I203_V1.5901.00
173_T185_E1.5861.00
37_L156_A1.5841.00
177_R180_G1.5741.00
14_N350_V1.5691.00
173_T194_K1.5691.00
37_L166_V1.5641.00
21_D25_R1.5641.00
468_W474_G1.5631.00
258_V266_S1.5571.00
467_L482_Q1.5411.00
284_I289_I1.5351.00
30_L167_L1.5211.00
410_T413_L1.5181.00
27_L376_I1.5151.00
431_N452_K1.5001.00
293_L306_R1.4901.00
258_V296_Y1.4821.00
432_A439_G1.4801.00
441_D445_E1.4801.00
167_L195_Y1.4741.00
176_R185_E1.4731.00
428_L449_A1.4711.00
213_F216_D1.4681.00
186_A195_Y1.4671.00
175_A213_F1.4651.00
440_E484_R1.4641.00
230_K275_Y1.4641.00
32_L197_W1.4591.00
27_L370_T1.4581.00
293_L297_N1.4571.00
306_R309_I1.4551.00
20_A155_N1.4481.00
155_N358_Y1.4381.00
348_L369_L1.4371.00
24_G162_K1.4241.00
86_R142_S1.4201.00
170_T195_Y1.4171.00
297_N303_Q1.4161.00
227_R321_D1.4131.00
267_I313_Y1.4101.00
231_G317_R1.4071.00
38_A152_V1.3911.00
232_S273_V1.3691.00
178_E183_I1.3691.00
158_M162_K1.3691.00
363_E382_K1.3681.00
40_A44_A1.3661.00
368_K371_P1.3631.00
92_R135_I1.3621.00
38_A149_A1.3571.00
187_E192_G1.3531.00
480_D484_R1.3531.00
172_A184_V1.3441.00
6_L22_A1.3421.00
427_L430_G1.3261.00
245_A249_Q1.3150.99
441_D447_Y1.3010.99
19_A164_G1.2990.99
151_L355_L1.2860.99
22_A366_L1.2810.99
11_G37_L1.2750.99
147_D420_T1.2670.99
167_L170_T1.2600.99
46_S259_I1.2570.99
201_G348_L1.2540.99
482_Q486_S1.2400.99
466_A489_L1.2390.99
210_V226_I1.2360.99
237_P264_E1.2300.99
403_R411_E1.2260.99
65_S69_A1.2210.99
296_Y304_L1.2140.99
217_G339_H1.2140.99
18_I362_A1.2040.99
208_P316_V1.2020.99
18_I205_A1.2000.99
5_D27_L1.1900.99
201_G373_Y1.1900.99
398_Y418_A1.1790.99
481_Q485_V1.1770.99
435_V438_L1.1730.99
398_Y426_E1.1650.99
86_R140_E1.1640.99
4_K28_S1.1610.99
370_T376_I1.1560.99
82_A144_C1.1480.99
238_R263_D1.1440.99
246_Y249_Q1.1410.99
74_L82_A1.1380.98
174_S216_D1.1350.98
19_A159_V1.1350.98
158_M161_R1.1340.98
362_A379_A1.1340.98
108_Y112_G1.1310.98
401_R405_R1.1290.98
21_D379_A1.1260.98
420_T450_E1.1220.98
62_R66_E1.1140.98
452_K456_D1.1100.98
60_E63_L1.1100.98
452_K488_W1.1090.98
251_E298_T1.1050.98
31_M37_L1.1050.98
88_R138_G1.1030.98
78_A386_L1.1030.98
319_L354_K1.0970.98
368_K372_Y1.0920.98
338_I348_L1.0910.98
442_F451_L1.0830.98
234_I268_I1.0820.98
66_E360_K1.0780.98
230_K285_E1.0710.98
188_D193_K1.0660.98
370_T373_Y1.0640.98
403_R414_A1.0620.98
205_A350_V1.0620.98
58_H329_A1.0600.98
271_T317_R1.0600.98
175_A216_D1.0590.98
336_L364_H1.0530.97
124_R127_A1.0530.97
485_V488_W1.0510.97
398_Y402_L1.0510.97
275_Y278_D1.0410.97
244_Q247_I1.0410.97
278_D281_A1.0390.97
83_F145_W1.0380.97
476_W482_Q1.0320.97
240_H308_D1.0320.97
28_S165_E1.0320.97
235_V311_W1.0250.97
7_I202_L1.0220.97
411_E415_R1.0210.97
21_D362_A1.0140.97
431_N456_D1.0130.97
453_Y457_H1.0090.97
178_E198_Q1.0090.97
390_A423_S1.0080.97
444_H473_Q1.0060.97
41_T208_P1.0020.97
48_L145_W1.0020.97
104_G109_D1.0010.97
348_L368_K0.9970.97
96_R99_W0.9870.96
32_L170_T0.9870.96
175_A181_L0.9840.96
133_P301_K0.9830.96
169_R461_R0.9800.96
231_G271_T0.9760.96
434_T437_D0.9710.96
183_I198_Q0.9670.96
477_L482_Q0.9640.96
417_Y428_L0.9600.96
338_I368_K0.9570.96
468_W476_W0.9480.95
178_E182_W0.9340.95
74_L151_L0.9330.95
66_E333_D0.9310.95
63_L67_A0.9310.95
124_R128_N0.9280.95
156_A168_T0.9270.95
55_Y354_K0.9250.95
4_K7_I0.9240.95
17_G362_A0.9230.95
172_A213_F0.9220.95
275_Y316_V0.9210.95
83_F419_R0.9170.94
80_H388_G0.9160.94
128_N131_L0.9120.94
14_N361_L0.9090.94
70_E359_R0.9050.94
467_L486_S0.9030.94
322_D330_I0.9000.94
350_V365_A0.9000.94
5_D28_S0.8990.94
175_A184_V0.8980.94
23_A29_V0.8980.94
35_Q40_A0.8970.94
477_L481_Q0.8960.94
161_R387_P0.8950.94
69_A360_K0.8940.94
199_A202_L0.8860.93
377_G380_W0.8860.93
438_L449_A0.8830.93
449_A452_K0.8830.93
184_V199_A0.8830.93
396_D399_A0.8820.93
385_V389_G0.8810.93
350_V361_L0.8810.93
54_R270_T0.8760.93
19_A166_V0.8730.93
183_I196_S0.8710.93
35_Q169_R0.8700.93
73_V77_M0.8680.93
237_P310_V0.8670.93
256_V295_V0.8630.92
123_L128_N0.8630.92
174_S194_K0.8620.92
170_T197_W0.8610.92
430_G434_T0.8610.92
201_G372_Y0.8580.92
49_I74_L0.8570.92
428_L432_A0.8530.92
397_D400_A0.8520.92
398_Y401_R0.8500.92
157_Q160_V0.8480.92
432_A449_A0.8450.92
415_R419_R0.8440.92
248_L258_V0.8440.92
132_K303_Q0.8410.91
369_L373_Y0.8410.91
481_Q484_R0.8410.91
363_E379_A0.8400.91
414_A425_S0.8390.91
424_N427_L0.8390.91
329_A335_T0.8350.91
64_V68_L0.8350.91
231_G315_G0.8330.91
102_R106_F0.8320.91
446_F450_E0.8310.91
211_K214_F0.8260.91
152_V355_L0.8240.91
85_M145_W0.8240.91
317_R332_R0.8240.91
373_Y376_I0.8230.91
183_I346_P0.8230.91
109_D112_G0.8230.91
52_G55_Y0.8200.90
190_D193_K0.8190.90
285_E288_E0.8190.90
399_A402_L0.8180.90
317_R354_K0.8170.90
401_R404_R0.8170.90
76_K383_E0.8170.90
336_L365_A0.8160.90
21_D367_E0.8160.90
320_C335_T0.8140.90
367_E381_T0.8140.90
430_G433_G0.8090.90
406_Y429_L0.8080.90
367_E379_A0.8080.90
432_A438_L0.8070.90
54_R354_K0.8060.90
271_T354_K0.8050.90
279_P282_V0.8010.89
431_N457_H0.7980.89
23_A162_K0.7970.89
243_K248_L0.7950.89
17_G21_D0.7920.89
31_M156_A0.7900.89
239_V302_K0.7870.89
8_V217_G0.7860.88
334_Y364_H0.7850.88
209_W279_P0.7840.88
37_L459_W0.7820.88
441_D481_Q0.7820.88
162_K384_S0.7820.88
187_E191_T0.7820.88
189_I193_K0.7760.88
66_E364_H0.7740.88
78_A81_I0.7730.88
8_V20_A0.7700.87
432_A436_S0.7680.87
400_A404_R0.7670.87
50_H54_R0.7670.87
431_N449_A0.7660.87
79_P389_G0.7660.87
22_A27_L0.7660.87
45_S148_D0.7660.87
122_G137_R0.7650.87
443_G484_R0.7650.87
363_E367_E0.7640.87
446_F471_T0.7640.87
380_W384_S0.7600.87
64_V108_Y0.7590.87
425_S429_L0.7550.86
436_S441_D0.7550.86
23_A164_G0.7540.86
409_L414_A0.7540.86
268_I292_L0.7510.86
212_Q279_P0.7500.86
319_L332_R0.7490.86
233_H271_T0.7480.86
154_A386_L0.7470.86
25_R367_E0.7470.86
284_I288_E0.7460.86
439_G442_F0.7440.86
227_R330_I0.7410.85
463_A467_L0.7410.85
402_L425_S0.7390.85
191_T194_K0.7390.85
189_I480_D0.7380.85
24_G158_M0.7370.85
123_L140_E0.7370.85
268_I296_Y0.7360.85
363_E381_T0.7360.85
233_H272_D0.7340.85
62_R333_D0.7310.85
22_A29_V0.7310.85
360_K364_H0.7290.84
482_Q485_V0.7280.84
106_F110_H0.7280.84
244_Q249_Q0.7280.84
317_R353_G0.7270.84
416_H450_E0.7210.84
211_K215_D0.7190.84
435_V448_E0.7190.84
452_K455_V0.7170.83
89_L101_I0.7160.83
29_V164_G0.7150.83
413_L447_Y0.7150.83
300_F303_Q0.7140.83
271_T318_P0.7130.83
348_L365_A0.7130.83
270_T354_K0.7120.83
92_R134_E0.7120.83
108_Y111_L0.7110.83
467_L489_L0.7110.83
273_V288_E0.7080.83
417_Y425_S0.7060.82
478_N484_R0.7060.82
294_N298_T0.7050.82
451_L489_L0.7050.82
326_S329_A0.7050.82
6_L366_L0.7030.82
271_T332_R0.7030.82
416_H446_F0.7020.82
230_K314_S0.7010.82
104_G111_L0.7000.82
204_N349_S0.6980.82
231_G354_K0.6950.81
417_Y447_Y0.6950.81
425_S428_L0.6950.81
318_P354_K0.6950.81
476_W481_Q0.6940.81
485_V489_L0.6940.81
262_M419_R0.6920.81
106_F120_S0.6910.81
20_A24_G0.6910.81
232_S284_I0.6900.81
159_V362_A0.6890.81
391_I423_S0.6870.81
252_D270_T0.6860.81
242_Q248_L0.6860.81
235_V313_Y0.6860.81
396_D400_A0.6840.80
274_E330_I0.6830.80
229_I319_L0.6830.80
64_V319_L0.6830.80
23_A159_V0.6820.80
297_N305_S0.6820.80
395_R398_Y0.6820.80
12_G15_G0.6820.80
190_D462_R0.6760.80
183_I199_A0.6750.80
92_R131_L0.6750.80
104_G107_M0.6750.80
10_G15_G0.6740.79
76_K115_T0.6710.79
254_R257_F0.6700.79
104_G108_Y0.6690.79
466_A485_V0.6680.79
210_V349_S0.6670.79
140_E144_C0.6660.79
60_E333_D0.6640.78
217_G345_A0.6640.78
402_L414_A0.6640.78
95_L98_A0.6640.78
20_A384_S0.6640.78
254_R354_K0.6640.78
81_I154_A0.6630.78
6_L29_V0.6610.78
479_A483_S0.6610.78
360_K363_E0.6610.78
9_I172_A0.6600.78
483_S487_Q0.6590.78
132_K135_I0.6580.78
415_R434_T0.6550.78
187_E193_K0.6550.78
291_Y300_F0.6550.78
49_I131_L0.6540.77
400_A411_E0.6540.77
41_T354_K0.6530.77
138_G247_I0.6520.77
478_N482_Q0.6510.77
8_V366_L0.6500.77
29_V166_V0.6500.77
257_F270_T0.6500.77
91_H95_L0.6490.77
434_T452_K0.6480.77
95_L101_I0.6480.77
287_S290_N0.6470.77
103_I107_M0.6460.77
439_G448_E0.6460.77
214_F347_L0.6460.77
75_L82_A0.6450.76
21_D363_E0.6450.76
184_V197_W0.6440.76
46_S257_F0.6430.76
435_V441_D0.6420.76
203_V366_L0.6410.76
443_G446_F0.6400.76
252_D257_F0.6400.76
481_Q486_S0.6370.76
212_Q215_D0.6370.76
181_L198_Q0.6360.75
321_D325_D0.6360.75
18_I365_A0.6360.75
302_K452_K0.6310.75
277_G281_A0.6300.75
332_R354_K0.6300.75
55_Y317_R0.6300.75
146_V355_L0.6280.75
413_L445_E0.6280.75
434_T456_D0.6260.74
129_S135_I0.6260.74
6_L371_P0.6230.74
381_T388_G0.6220.74
271_T319_L0.6210.74
235_V267_I0.6210.74
210_V351_F0.6210.74
404_R410_T0.6200.74
51_G56_L0.6200.74
272_D354_K0.6190.73
50_H354_K0.6190.73
159_V166_V0.6180.73
30_L186_A0.6180.73
251_E294_N0.6150.73
25_R379_A0.6140.73
25_R381_T0.6130.73
410_T414_A0.6130.73
47_K148_D0.6130.73
446_F451_L0.6110.73
50_H55_Y0.6110.73
55_Y231_G0.6100.72
339_H344_K0.6100.72
91_H101_I0.6090.72
56_L64_V0.6080.72
55_Y319_L0.6070.72
243_K246_Y0.6070.72
176_R198_Q0.6060.72
183_I202_L0.6060.72
142_S145_W0.6060.72
480_D483_S0.6050.72
432_A435_V0.6030.72
428_L447_Y0.6030.72
93_P98_A0.6020.71
204_N210_V0.6020.71
233_H254_R0.6010.71
397_D404_R0.5990.71
181_L347_L0.5980.71
455_V460_V0.5980.71
6_L18_I0.5960.71
44_A313_Y0.5960.71
140_E143_D0.5960.71
402_L426_E0.5950.71
176_R181_L0.5930.70
359_R381_T0.5920.70
420_T473_Q0.5920.70
14_N362_A0.5900.70
437_D440_E0.5900.70
405_R433_G0.5880.70
219_H222_S0.5880.70
430_G457_H0.5870.70
436_S440_E0.5870.70
366_L379_A0.5870.70
247_I255_I0.5870.70
27_L374_Q0.5850.69
42_S148_D0.5850.69
191_T376_I0.5850.69
231_G319_L0.5830.69
86_R91_H0.5830.69
84_P142_S0.5820.69
426_E430_G0.5810.69
30_L165_E0.5800.69
170_T173_T0.5790.68
91_H139_F0.5780.68
77_M358_Y0.5780.68
484_R488_W0.5780.68
193_K440_E0.5770.68
401_R426_E0.5760.68
101_I104_G0.5760.68
446_F454_L0.5760.68
76_K116_S0.5750.68
240_H258_V0.5740.68
171_R212_Q0.5730.68
446_F466_A0.5720.68
232_S312_T0.5720.68
358_Y381_T0.5720.68
121_T127_A0.5720.68
38_A43_S0.5700.67
451_L460_V0.5670.67
441_D444_H0.5660.67
189_I192_G0.5660.67
234_I328_Q0.5650.67
396_D415_R0.5650.67
273_V285_E0.5650.67
273_V287_S0.5650.67
191_T461_R0.5650.67
463_A489_L0.5640.66
189_I194_K0.5620.66
326_S330_I0.5600.66
432_A437_D0.5600.66
8_V29_V0.5600.66
215_D224_Y0.5600.66
272_D317_R0.5590.66
156_A160_V0.5580.66
69_A363_E0.5580.66
269_G315_G0.5580.66
455_V463_A0.5570.66
400_A407_P0.5570.66
52_G354_K0.5560.65
286_E289_I0.5540.65
4_K186_A0.5540.65
65_S382_K0.5540.65
473_Q485_V0.5510.65
58_H273_V0.5510.65
50_H356_T0.5500.65
399_A415_R0.5490.64
409_L413_L0.5490.64
428_L431_N0.5480.64
176_R179_N0.5470.64
432_A447_Y0.5470.64
160_V459_W0.5460.64
236_V304_L0.5460.64
438_L442_F0.5450.64
108_Y141_Y0.5430.64
72_E84_P0.5430.64
435_V439_G0.5430.64
212_Q216_D0.5430.64
88_R123_L0.5400.63
105_L108_Y0.5400.63
229_I272_D0.5390.63
433_G436_S0.5380.63
464_D468_W0.5370.63
182_W198_Q0.5370.63
438_L448_E0.5360.63
463_A486_S0.5350.62
51_G356_T0.5350.62
157_Q161_R0.5340.62
400_A415_R0.5340.62
78_A359_R0.5310.62
334_Y361_L0.5310.62
369_L374_Q0.5310.62
172_A216_D0.5300.62
30_L34_A0.5300.62
234_I289_I0.5300.62
16_A166_V0.5290.62
363_E380_W0.5290.62
51_G55_Y0.5290.62
322_D325_D0.5280.61
454_L457_H0.5280.61
395_R399_A0.5270.61
185_E195_Y0.5270.61
391_I398_Y0.5260.61
442_F448_E0.5260.61
278_D282_V0.5260.61
20_A162_K0.5260.61
16_A31_M0.5250.61
354_K357_T0.5250.61
473_Q481_Q0.5240.61
367_E370_T0.5240.61
443_G485_V0.5230.61
110_H113_K0.5230.61
52_G319_L0.5230.61
358_Y386_L0.5220.61
50_H317_R0.5220.61
80_H389_G0.5200.60
58_H102_R0.5190.60
47_K54_R0.5190.60
463_A466_A0.5190.60
455_V462_R0.5180.60
162_K380_W0.5180.60
175_A183_I0.5180.60
8_V371_P0.5160.60
295_V298_T0.5160.60
376_I380_W0.5150.60
70_E78_A0.5150.60
43_S148_D0.5140.59
329_A333_D0.5140.59
229_I327_P0.5130.59
112_G211_K0.5130.59
112_G120_S0.5130.59
64_V71_R0.5130.59
389_G392_E0.5120.59
275_Y282_V0.5120.59
21_D359_R0.5120.59
232_S288_E0.5120.59
92_R327_P0.5110.59
238_R241_T0.5110.59
429_L432_A0.5110.59
447_Y450_E0.5100.59
455_V486_S0.5090.59
332_R353_G0.5090.59
102_R120_S0.5090.59
158_M387_P0.5090.59
413_L417_Y0.5080.59
444_H476_W0.5080.59
438_L452_K0.5070.58
233_H319_L0.5070.58
337_D351_F0.5060.58
242_Q245_A0.5060.58
67_A357_T0.5050.58
108_Y252_D0.5040.58
337_D349_S0.5040.58
434_T438_L0.5040.58
49_I78_A0.5040.58
440_E488_W0.5040.58
13_I53_L0.5030.58
209_W230_K0.5030.58
229_I288_E0.5030.58
406_Y414_A0.5020.58
366_L369_L0.5020.58
68_L261_W0.5020.58
178_E185_E0.5010.57
232_S309_I0.5010.57
63_L68_L0.5010.57
67_A255_I0.5010.57
417_Y450_E0.5010.57
81_I355_L0.5010.57
451_L467_L0.5010.57
57_E64_V0.5000.57
261_W267_I0.5000.57
175_A347_L0.5000.57
Legend: The i and j are positions as given in the UniProt sequence. Only predictions with Prob ≥ 0.7 are listed. For all 3L/2 predictions, click on one of the links below.

Sco = ((raw_sco - min_sco)/(avg_sco - min_sco))/2 + 0.5
Prob = P(contact | Sco, seq80/√len)
seq80 = effective number of sequences, after reducing redundancy to 80% identity

HHsearch Results:
Legend: GREMLIN results (Prob ≥ 0.7) overlaid on top PDB hits. The grey/red filled circles underneath are pdb residue contacts (min distance < 5 Angstroms). The shade of the circles is based on the top 10 (or top 20 if e-value < 1E-20) HHsearch results which uses the overall probability and per-site alignment prob. Inter oligomeric contacts in the pdb are in shades of red.
PDB Chains Cov % HHΔ  
2qcuA 2 0.99 100 0.286 Contact Map
2rghA 2 0.9541 100 0.308 Contact Map
3da1A 1 0.8782 100 0.327 Contact Map
4h1bA 1 0.7026 100 0.628 Contact Map
4x9mA 1 0.7226 100 0.63 Contact Map
3dmeA 2 0.7006 100 0.637 Contact Map
3axbA 2 0.7246 100 0.649 Contact Map
1y56B 1 0.7186 100 0.651 Contact Map
1ryiA 4 0.6846 100 0.651 Contact Map
1c0pA 2 0.6467 100 0.653 Contact Map

Sequence Conservation (Image Generated using WebLogo v3)
Legend: Color is used to show Hydrophobicity (Blue: Hydrophilic, Green: Neutral and Black: Hydrophobic). Amount of gaps is represented by width.

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