GREMLIN Database
CARY - Carbamoyl-phosphate synthase arginine-specific large chain
UniProt: P18185 Sequences:
Download Alignment
We filter this alignment to remove sites that have many gaps before running GREMLIN.

Date: 2015_01
Length: 1030 (1021)
Sequences: 2856 (1138)
Seq/√Len: 35.6

GREMLIN Results:

Legend: The darker and larger the blue dots, the higher coevolution strength.

Residue pairs sorted by coevolution strength:
i j Sco Prob
355_V392_A7.0691.00
463_D467_K5.4001.00
226_N273_S4.8321.00
9_S43_R4.6241.00
9_S45_I4.5951.00
559_L595_I4.3251.00
69_L98_L4.2931.00
288_D291_S4.2501.00
702_S727_Q4.2361.00
104_E124_Q4.2181.00
680_E812_K4.1701.00
82_R111_H4.1621.00
860_R865_A4.0911.00
663_Q673_Q4.0601.00
556_K632_D3.9041.00
155_A192_I3.6571.00
192_I204_C3.6001.00
619_T623_V3.5631.00
936_L950_A3.5551.00
437_E441_R3.5161.00
933_S960_A3.4971.00
218_Y233_N3.4341.00
288_D293_Q3.4271.00
398_A621_H3.3931.00
913_A918_S3.3481.00
10_I42_Y3.3281.00
138_E274_A3.2911.00
792_P880_V3.2361.00
804_D808_K3.1341.00
237_I462_F3.1001.00
764_E866_S3.0851.00
78_I111_H3.0661.00
233_N253_S3.0651.00
264_L294_Y3.0531.00
435_V448_I3.0271.00
75_T110_E3.0211.00
145_E208_K2.9781.00
79_K111_H2.9721.00
708_V746_I2.9711.00
595_I625_V2.9691.00
439_L459_L2.9601.00
468_L490_K2.8851.00
866_S869_D2.8761.00
42_Y327_V2.8401.00
513_K627_E2.8231.00
75_T79_K2.7801.00
156_L189_T2.7771.00
160_E185_K2.7501.00
940_P943_V2.7241.00
865_A869_D2.7081.00
43_R63_E2.7061.00
698_A731_S2.6941.00
759_L806_A2.6761.00
760_I861_L2.6591.00
446_D456_P2.6501.00
796_S800_Q2.6291.00
75_T111_H2.6151.00
4_D114_K2.5781.00
970_L974_L2.5761.00
546_T628_Q2.5691.00
559_L626_A2.5461.00
130_E134_S2.5341.00
689_E703_E2.5181.00
393_I432_F2.5151.00
149_V158_F2.5121.00
884_F923_I2.5101.00
390_E461_T2.4961.00
439_L445_I2.4911.00
185_K189_T2.4521.00
481_F485_K2.4511.00
955_H977_S2.4461.00
10_I327_V2.4421.00
706_Y727_Q2.4381.00
421_W447_D2.4071.00
74_L98_L2.3981.00
386_E391_A2.3961.00
555_K591_E2.3871.00
968_N978_E2.3831.00
159_A164_F2.3711.00
684_K805_A2.3711.00
47_V77_I2.3400.99
485_K489_E2.3220.99
587_E856_P2.2890.99
71_A101_E2.2860.99
272_I283_I2.2840.99
471_R486_K2.2810.99
151_N154_D2.2790.99
933_S962_W2.2750.99
626_A655_I2.2750.99
726_S729_Q2.2520.99
799_Q876_N2.2460.99
218_Y253_S2.2450.99
75_T105_T2.2350.99
416_S419_T2.2300.99
689_E700_K2.2230.99
866_S870_L2.2080.99
664_T814_S2.1920.99
141_Q270_T2.1910.99
857_L870_L2.1880.99
559_L625_V2.1850.99
509_R623_V2.1470.99
39_E324_K2.1450.99
553_K591_E2.1440.99
807_Q811_R2.1350.99
504_T507_E2.1280.99
443_V447_D2.1250.99
242_V427_P2.1180.99
208_K293_Q2.1140.99
142_P293_Q2.1040.99
483_L487_A2.1040.99
271_I296_V2.0900.99
164_F185_K2.0860.99
150_D203_Q2.0830.99
683_L805_A2.0820.99
769_P854_M2.0500.99
370_R429_D2.0490.99
765_E868_K2.0460.99
375_K891_A2.0430.99
100_V120_V2.0400.99
801_G804_D2.0380.99
947_A970_L2.0180.98
394_Q492_F2.0170.98
475_A480_S2.0080.98
439_L448_I2.0070.98
164_F184_D2.0030.98
506_E652_K1.9930.98
71_A105_T1.9870.98
390_E464_N1.9810.98
960_A979_E1.9800.98
755_V794_I1.9770.98
141_Q267_A1.9760.98
417_I441_R1.9650.98
147_E161_S1.9620.98
737_E744_I1.9580.98
651_E663_Q1.9470.98
698_A727_Q1.9380.98
76_A79_K1.9360.98
171_A203_Q1.9300.98
666_D714_Y1.9070.98
666_D673_Q1.8990.98
613_Y629_E1.8980.98
727_Q731_S1.8970.98
79_K110_E1.8970.98
513_K518_T1.8880.98
287_L294_Y1.8820.98
75_T107_V1.8800.97
78_I113_V1.8770.97
40_E324_K1.8760.97
182_A191_M1.8760.97
159_A166_V1.8720.97
66_F910_M1.8540.97
445_I459_L1.8510.97
587_E860_R1.8370.97
797_G828_G1.8330.97
693_T696_E1.8270.97
47_V74_L1.8270.97
435_V454_I1.8220.97
757_I806_A1.8180.97
253_S261_Y1.8160.97
511_L515_M1.8140.97
699_S703_E1.8110.97
687_K703_E1.8100.97
648_E652_K1.8030.97
103_E120_V1.7990.97
968_N980_A1.7980.97
929_S932_G1.7830.97
184_D187_A1.7810.97
981_R995_F1.7810.97
588_L860_R1.7780.96
947_A974_L1.7710.96
691_A700_K1.7710.96
150_D154_D1.7660.96
494_D512_R1.7520.96
470_N473_M1.7470.96
942_D1007_S1.7460.96
9_S63_E1.7400.96
558_A658_L1.7340.96
147_E158_F1.7280.96
755_V825_I1.7210.96
684_K829_N1.7180.96
732_Q736_D1.7090.96
1004_F1024_E1.7020.95
755_V802_M1.6980.95
738_D744_I1.6960.95
151_N202_N1.6930.95
436_M440_S1.6920.95
135_L274_A1.6840.95
720_G737_E1.6820.95
518_T627_E1.6790.95
761_S811_R1.6750.95
439_L462_F1.6730.95
168_I204_C1.6710.95
677_L809_I1.6660.95
436_M462_F1.6660.95
795_T828_G1.6630.95
444_S447_D1.6580.95
168_I206_V1.6470.95
801_G805_A1.6440.95
449_H459_L1.6420.94
424_A438_L1.6380.94
124_Q131_K1.6330.94
183_P187_A1.6250.94
795_T798_L1.6120.94
510_A514_E1.6110.94
196_L202_N1.6090.94
990_H994_V1.6090.94
275_L299_V1.6060.94
533_A536_N1.6010.94
388_N543_F1.5930.94
260_D292_K1.5890.93
588_L863_A1.5830.93
619_T649_A1.5810.93
164_F188_F1.5780.93
687_K748_Q1.5720.93
962_W1001_V1.5710.93
125_K274_A1.5610.93
759_L819_M1.5570.93
465_I492_F1.5520.93
677_L812_K1.5490.93
382_V399_S1.5480.93
963_M978_E1.5470.92
133_R137_N1.5460.92
103_E109_K1.5440.92
71_A74_L1.5390.92
389_L457_F1.5380.92
73_S77_I1.5360.92
168_I188_F1.5350.92
1022_K1026_A1.5310.92
585_L863_A1.5300.92
798_L828_G1.5290.92
755_V830_I1.5280.92
684_K801_G1.5270.92
218_Y268_S1.5240.92
585_L592_T1.5230.92
224_S330_T1.5200.92
397_A432_F1.5170.92
71_A98_L1.5160.92
480_S483_L1.5010.91
556_K590_F1.5000.91
464_N490_K1.4960.91
915_D918_S1.4960.91
675_Y685_H1.4940.91
798_L830_I1.4920.91
791_G799_Q1.4900.91
149_V154_D1.4870.91
825_I830_I1.4850.91
86_L113_V1.4800.91
523_I538_F1.4790.91
801_G830_I1.4760.91
508_V511_L1.4720.90
107_V111_H1.4680.90
511_L514_E1.4670.90
405_I499_S1.4650.90
443_V448_I1.4640.90
983_E1016_S1.4640.90
619_T653_A1.4600.90
393_I407_T1.4580.90
484_L511_L1.4520.90
786_F884_F1.4500.90
760_I854_M1.4440.90
635_I859_T1.4440.90
505_E509_R1.4390.89
497_I501_I1.4390.89
88_A99_A1.4370.89
144_P295_Y1.4310.89
798_L825_I1.4300.89
468_L487_A1.4300.89
805_A830_I1.4250.89
859_T863_A1.4240.89
800_Q803_K1.4240.89
767_F872_P1.4220.89
632_D656_T1.4220.89
673_Q714_Y1.4010.88
421_W443_V1.3990.88
664_T667_V1.3950.88
865_A870_L1.3930.88
938_N946_L1.3910.88
149_V155_A1.3900.88
558_A863_A1.3830.87
678_L832_V1.3740.87
225_N330_T1.3740.87
421_W451_K1.3720.87
449_H453_K1.3650.87
321_M337_P1.3620.87
592_T863_A1.3610.87
513_K624_N1.3590.87
152_E188_F1.3570.86
683_L830_I1.3550.86
237_I248_I1.3540.86
420_L448_I1.3520.86
472_L487_A1.3520.86
69_L74_L1.3500.86
750_V826_D1.3470.86
928_W932_G1.3460.86
134_S138_E1.3450.86
101_E104_E1.3420.86
806_A823_F1.3410.86
495_A505_E1.3350.86
667_V812_K1.3340.85
950_A961_S1.3260.85
159_A188_F1.3230.85
468_L483_L1.3220.85
634_A655_I1.3180.85
701_A706_Y1.3140.85
553_K557_R1.3130.85
439_L463_D1.3060.84
1018_K1022_K1.3050.84
1008_G1014_P1.3040.84
558_A585_L1.2950.84
761_S766_V1.2930.84
557_R591_E1.2910.84
472_L500_L1.2890.83
465_I490_K1.2820.83
481_F515_M1.2810.83
417_I447_D1.2790.83
6_S114_K1.2780.83
221_M321_M1.2770.83
70_T73_S1.2770.83
801_G829_N1.2760.83
263_M287_L1.2750.83
237_I435_V1.2740.83
969_S973_N1.2670.82
518_T624_N1.2670.82
257_T260_D1.2660.82
972_I978_E1.2630.82
40_E329_Y1.2540.82
341_S534_K1.2530.82
394_Q406_G1.2530.82
972_I980_A1.2530.82
982_K991_G1.2530.82
152_E156_L1.2500.81
809_I821_I1.2490.81
461_T464_N1.2490.81
237_I250_V1.2480.81
126_G283_I1.2450.81
394_Q465_I1.2420.81
146_S207_E1.2410.81
11_L86_L1.2320.81
100_V131_K1.2310.80
475_A483_L1.2260.80
440_S469_E1.2240.80
393_I462_F1.2230.80
479_L484_L1.2200.80
404_N433_F1.2150.80
623_V655_I1.2090.79
635_I862_L1.2070.79
734_L737_E1.1980.79
457_F461_T1.1980.79
675_Y688_G1.1970.79
329_Y334_L1.1960.78
734_L738_D1.1890.78
443_V451_K1.1860.78
472_L497_I1.1860.78
676_Q680_E1.1840.78
194_Q198_A1.1830.78
157_H160_E1.1780.77
503_K508_V1.1780.77
788_I916_S1.1770.77
479_L501_I1.1710.77
965_Q980_A1.1640.77
503_K507_E1.1620.76
799_Q803_K1.1570.76
1008_G1017_L1.1570.76
583_L608_I1.1510.76
550_I628_Q1.1500.76
125_K135_L1.1490.76
549_D552_R1.1490.76
936_L971_H1.1440.75
170_P179_G1.1400.75
71_A107_V1.1400.75
771_Y803_K1.1390.75
86_L108_L1.1370.75
126_G720_G1.1370.75
588_L865_A1.1360.75
184_D989_T1.1350.75
424_A452_T1.1350.75
155_A206_V1.1340.75
519_P542_Y1.1290.74
611_R629_E1.1270.74
47_V102_L1.1260.74
405_I495_A1.1250.74
71_A75_T1.1240.74
34_C60_F1.1210.74
766_V811_R1.1150.73
131_K134_S1.1140.73
267_A294_Y1.1140.73
720_G738_D1.1130.73
436_M466_I1.1130.73
561_I622_I1.1100.73
250_V355_V1.1080.73
125_K138_E1.1080.73
124_Q172_Y1.1060.73
805_A823_F1.1060.73
208_K295_Y1.1050.73
99_A115_L1.1040.73
448_I462_F1.1040.73
390_E490_K1.1030.73
423_L434_V1.1000.72
449_H456_P1.1000.72
13_I74_L1.1000.72
417_I421_W1.0970.72
133_R207_E1.0920.72
861_L870_L1.0910.72
471_R483_L1.0880.72
621_H624_N1.0840.71
13_I95_A1.0830.71
632_D658_L1.0830.71
221_M272_I1.0800.71
522_K545_E1.0790.71
724_V744_I1.0780.71
845_V910_M1.0770.71
337_P606_Y1.0740.71
227_T345_S1.0720.70
100_V124_Q1.0720.70
423_L428_D1.0720.70
952_F956_E1.0690.70
218_Y228_C1.0670.70
471_R475_A1.0650.70
388_N490_K1.0650.70
321_M334_L1.0650.70
188_F206_V1.0650.70
989_T994_V1.0650.70
938_N950_A1.0570.69
984_R993_P1.0560.69
437_E440_S1.0520.69
141_Q296_V1.0520.69
509_R653_A1.0510.69
155_A168_I1.0470.69
683_L801_G1.0470.69
137_N143_V1.0460.68
552_R611_R1.0460.68
340_G534_K1.0450.68
142_P208_K1.0440.68
771_Y799_Q1.0440.68
258_D262_Q1.0430.68
262_Q265_R1.0400.68
484_L517_I1.0390.68
228_C269_L1.0380.68
191_M195_A1.0370.68
809_I832_V1.0350.68
13_I69_L1.0340.68
382_V396_A1.0310.67
228_C273_S1.0300.67
259_E292_K1.0290.67
806_A821_I1.0280.67
582_V609_A1.0280.67
250_V389_L1.0270.67
934_I1005_L1.0260.67
623_V653_A1.0240.67
227_T330_T1.0220.67
121_E131_K1.0220.67
19_I177_K1.0220.67
438_L443_V1.0210.67
498_A504_T1.0180.66
36_A334_L1.0170.66
336_N339_T1.0170.66
214_K455_D1.0160.66
207_E210_I1.0160.66
624_N627_E1.0150.66
983_E1020_L1.0150.66
141_Q283_I1.0150.66
192_I206_V1.0140.66
388_N392_A1.0110.66
168_I192_I1.0100.66
420_L441_R1.0090.66
996_T1020_L1.0050.65
278_V325_L1.0000.65
266_T270_T1.0000.65
38_K60_F1.0000.65
159_A192_I0.9990.65
789_L846_V0.9980.65
937_Q967_G0.9930.64
617_M646_A0.9900.64
448_I459_L0.9900.64
962_W979_E0.9880.64
982_K990_H0.9870.64
101_E105_T0.9860.64
35_I607_E0.9820.64
731_S735_N0.9810.63
860_R870_L0.9810.63
228_C272_I0.9810.63
357_F360_W0.9790.63
290_F453_K0.9770.63
582_V592_T0.9760.63
697_A734_L0.9740.63
424_A434_V0.9740.63
786_F885_P0.9730.63
811_R814_S0.9720.63
714_Y742_Y0.9710.63
996_T1023_K0.9710.63
121_E125_K0.9700.63
849_V881_A0.9680.62
1_M7_I0.9630.62
961_S978_E0.9620.62
85_G88_A0.9610.62
933_S979_E0.9610.62
387_R522_K0.9600.62
692_Y696_E0.9600.62
225_N332_D0.9590.62
262_Q530_E0.9560.61
271_I631_I0.9550.61
234_M249_V0.9530.61
881_A912_V0.9510.61
816_K861_L0.9500.61
935_Y964_E0.9500.61
189_T193_K0.9430.60
184_D998_E0.9430.60
218_Y261_Y0.9420.60
404_N410_P0.9410.60
262_Q531_F0.9400.60
107_V110_E0.9390.60
408_H499_S0.9390.60
164_F189_T0.9380.60
145_E293_Q0.9380.60
982_K988_M0.9370.60
579_V1018_K0.9350.60
932_G959_F0.9350.60
221_M229_I0.9340.60
353_V392_A0.9340.60
791_G794_I0.9330.59
522_K543_F0.9300.59
253_S350_L0.9280.59
460_H463_D0.9270.59
445_I463_D0.9250.59
133_R148_I0.9230.59
468_L471_R0.9220.59
220_V230_T0.9220.59
1016_S1020_L0.9200.58
417_I443_V0.9170.58
213_F295_Y0.9120.58
462_F466_I0.9120.58
24_A566_I0.9110.58
694_K731_S0.9100.58
757_I802_M0.9090.57
320_K337_P0.9090.57
394_Q457_F0.9080.57
4_D42_Y0.9080.57
519_P624_N0.9070.57
591_E611_R0.9060.57
180_G191_M0.9060.57
45_I77_I0.9050.57
773_E793_S0.9030.57
144_P210_I0.9030.57
581_G635_I0.9020.57
638_F858_A0.9020.57
8_S43_R0.9000.57
481_F517_I0.8980.57
936_L961_S0.8980.57
268_S285_F0.8950.56
933_S995_F0.8940.56
156_L168_I0.8940.56
468_L486_K0.8940.56
964_E967_G0.8930.56
752_G824_V0.8920.56
734_L744_I0.8910.56
766_V807_Q0.8900.56
498_A508_V0.8890.56
379_T566_I0.8890.56
738_D742_Y0.8880.56
258_D531_F0.8860.56
559_L632_D0.8850.55
879_G912_V0.8840.55
210_I240_V0.8820.55
387_R543_F0.8820.55
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204_C211_A0.5280.27
468_L472_L0.5280.27
1006_Q1018_K0.5270.26
45_I63_E0.5270.26
82_R663_Q0.5260.26
237_I254_Q0.5260.26
435_V439_L0.5250.26
691_A755_V0.5250.26
67_E108_L0.5250.26
552_R555_K0.5250.26
72_E731_S0.5240.26
171_A179_G0.5240.26
850_M875_Q0.5240.26
585_L635_I0.5240.26
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795_T827_N0.5230.26
595_I1002_R0.5230.26
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702_S811_R0.5220.26
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126_G664_T0.5210.26
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827_N974_L0.5210.26
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382_V400_L0.5200.26
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213_F288_D0.5170.26
446_D635_I0.5170.26
336_N533_A0.5170.26
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550_I553_K0.5160.26
263_M291_S0.5160.26
278_V281_C0.5160.26
608_I833_L0.5160.26
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1008_G1012_P0.5150.26
152_E206_V0.5150.26
179_G203_Q0.5140.26
566_I688_G0.5140.26
855_I918_S0.5140.26
11_L115_L0.5140.26
314_T778_A0.5140.26
706_Y709_L0.5130.26
140_K153_A0.5130.26
133_R487_A0.5130.26
737_E909_G0.5130.26
509_R516_G0.5120.25
803_K875_Q0.5120.25
444_S449_H0.5120.25
337_P343_Y0.5120.25
196_L932_G0.5120.25
121_E124_Q0.5110.25
527_C536_N0.5110.25
495_A541_T0.5110.25
519_P825_I0.5110.25
963_M966_E0.5110.25
361_P885_P0.5110.25
419_T422_D0.5110.25
965_Q982_K0.5110.25
261_Y347_E0.5100.25
771_Y802_M0.5100.25
338_L536_N0.5100.25
1006_Q1016_S0.5090.25
739_S742_Y0.5090.25
471_R625_V0.5090.25
110_E931_K0.5090.25
808_K811_R0.5090.25
90_L523_I0.5090.25
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662_F846_V0.5080.25
82_R800_Q0.5080.25
996_T1004_F0.5080.25
525_D530_E0.5080.25
584_T588_L0.5080.25
480_S511_L0.5070.25
75_T86_L0.5070.25
617_M621_H0.5060.25
166_V207_E0.5060.25
44_V524_V0.5060.25
538_F788_I0.5060.25
882_V919_A0.5060.25
261_Y530_E0.5060.25
846_V854_M0.5060.25
142_P370_R0.5050.25
168_I961_S0.5050.25
983_E991_G0.5050.25
242_V452_T0.5050.25
83_P833_L0.5050.25
741_P744_I0.5050.25
452_T830_I0.5050.25
645_N788_I0.5040.25
4_D327_V0.5040.25
724_V971_H0.5040.25
422_D444_S0.5040.25
439_L970_L0.5030.25
10_I213_F0.5030.25
24_A597_N0.5030.25
141_Q363_D0.5030.25
552_R593_I0.5020.25
897_V1018_K0.5020.25
7_I768_I0.5020.25
204_C428_D0.5020.25
786_F923_I0.5020.25
571_G717_G0.5010.25
623_V634_A0.5010.25
194_Q619_T0.5010.25
181_I209_S0.5010.25
172_Y228_C0.5010.25
340_G465_I0.5010.25
340_G977_S0.5010.25
626_A650_L0.5010.25
10_I444_S0.5000.25
389_L393_I0.5000.25
3_K7_I0.5000.25
Legend: The i and j are positions as given in the UniProt sequence. Only predictions with Prob ≥ 0.7 are listed. For all 3L/2 predictions, click on one of the links below.

Sco = ((raw_sco - min_sco)/(avg_sco - min_sco))/2 + 0.5
Prob = P(contact | Sco, seq80/√len)
seq80 = effective number of sequences, after reducing redundancy to 80% identity

HHsearch Results:
Legend: GREMLIN results (Prob ≥ 0.7) overlaid on top PDB hits. The grey/red filled circles underneath are pdb residue contacts (min distance < 5 Angstroms). The shade of the circles is based on the top 10 (or top 20 if e-value < 1E-20) HHsearch results which uses the overall probability and per-site alignment prob. Inter oligomeric contacts in the pdb are in shades of red.
PDB Chains Cov % HHΔ  
1a9xA 3 0.9835 100 0.41 Contact Map
3jrxA 1 0.399 100 0.894 Contact Map
3glkA 1 0.3777 100 0.895 Contact Map
3orqA 2 0.3505 100 0.896 Contact Map
2vpqA 2 0.4019 100 0.898 Contact Map
1ulzA 2 0.4078 100 0.898 Contact Map
2dzdA 2 0.4019 100 0.901 Contact Map
1w96A 1 0.4049 100 0.902 Contact Map
3q2oA 2 0.3515 100 0.903 Contact Map
2dwcA 4 0.3621 100 0.905 Contact Map

Sequence Conservation (Image Generated using WebLogo v3)
Legend: Color is used to show Hydrophobicity (Blue: Hydrophilic, Green: Neutral and Black: Hydrophobic). Amount of gaps is represented by width.

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