GREMLIN Database
YUFO - Uncharacterized ABC transporter ATP-binding protein YufO
UniProt: O05253 Sequences:
Download Alignment
We filter this alignment to remove sites that have many gaps before running GREMLIN.

Date: 2015_01
Length: 510 (477)
Sequences: 9832 (6530)
Seq/√Len: 299.0

GREMLIN Results:

Legend: The darker and larger the blue dots, the higher coevolution strength.

Residue pairs sorted by coevolution strength:
i j Sco Prob
60_E67_K3.4501.00
21_D217_K3.4441.00
31_E209_R3.3151.00
33_H211_T3.1361.00
118_G135_E3.1201.00
317_N421_D3.1131.00
258_A279_S3.0961.00
313_T320_Q3.0551.00
14_F54_Y3.0501.00
385_I395_E3.0391.00
64_R160_D2.9971.00
301_E307_R2.8481.00
258_A315_T2.8171.00
9_N22_N2.7991.00
34_A207_C2.7771.00
261_G275_D2.7601.00
260_D313_T2.7531.00
116_R120_E2.7191.00
220_K235_L2.7091.00
122_Q126_D2.6421.00
136_A140_D2.5531.00
255_E258_A2.5401.00
13_A18_V2.5381.00
6_E62_R2.5101.00
6_E26_Q2.4771.00
300_I457_L2.4621.00
184_K205_E2.4401.00
254_A282_A2.4081.00
8_L60_E2.3961.00
385_I392_T2.3331.00
223_D226_D2.2861.00
207_C224_V2.2731.00
7_M10_I2.2361.00
120_E123_D2.2211.00
255_E279_S2.2131.00
299_L474_I2.2041.00
383_S387_E2.2031.00
127_R189_E2.1781.00
18_V21_D2.1421.00
70_I79_L2.1181.00
32_I183_M2.0791.00
347_V358_L2.0761.00
472_A482_S2.0751.00
11_R18_V2.0651.00
260_D315_T2.0301.00
177_K205_E2.0251.00
259_I262_I2.0171.00
234_S240_E2.0101.00
446_E450_A1.9651.00
274_R478_R1.9581.00
113_D116_R1.9491.00
456_L467_L1.9391.00
286_V472_A1.9341.00
47_M196_I1.9271.00
71_N75_K1.9121.00
384_L413_G1.9051.00
228_N231_E1.8961.00
9_N24_N1.8351.00
421_D452_K1.8291.00
8_L62_R1.8231.00
255_E281_K1.8011.00
319_K332_S1.7981.00
280_V453_A1.7841.00
289_A464_I1.7761.00
47_M83_M1.7431.00
259_I278_L1.7261.00
91_V102_L1.7211.00
11_R22_N1.7191.00
425_A444_L1.6961.00
256_V453_A1.6781.00
90_L138_A1.6741.00
382_R386_T1.6691.00
62_R67_K1.6591.00
316_L321_I1.6551.00
357_L377_M1.6551.00
184_K187_V1.6491.00
285_I444_L1.6471.00
287_G468_S1.6461.00
222_L235_L1.6351.00
432_L440_V1.6331.00
300_I424_I1.6321.00
84_V164_F1.6231.00
373_H376_E1.5951.00
185_N189_E1.5851.00
353_G396_Y1.5821.00
382_R395_E1.5751.00
211_T218_G1.5691.00
63_V161_I1.5581.00
399_A410_A1.5551.00
137_K140_D1.5521.00
352_I392_T1.5451.00
4_V192_S1.5411.00
10_I49_V1.5381.00
263_T275_D1.5331.00
13_A58_R1.5271.00
98_E114_R1.5231.00
397_T401_A1.5191.00
97_A118_G1.5161.00
160_D191_K1.5121.00
340_D343_K1.5111.00
93_T139_A1.5101.00
392_T397_T1.5091.00
123_D127_R1.5081.00
119_Q123_D1.5061.00
301_E344_H1.5011.00
256_V317_N1.4981.00
349_D400_R1.4891.00
316_L422_L1.4881.00
484_N487_E1.4811.00
222_L227_T1.4791.00
315_T320_Q1.4781.00
472_A479_I1.4691.00
385_I390_V1.4611.00
7_M50_L1.4481.00
280_V455_L1.4441.00
287_G464_I1.4381.00
116_R119_Q1.4371.00
127_R185_N1.4331.00
204_M232_L1.4291.00
66_E79_L1.4161.00
29_K192_S1.4141.00
387_E446_E1.4131.00
187_V208_D1.4101.00
75_K78_D1.4101.00
324_L328_K1.4061.00
446_E449_D1.4021.00
280_V422_L1.4001.00
302_A309_T1.4001.00
372_L377_M1.4001.00
356_I390_V1.3931.00
255_E282_A1.3911.00
72_S75_K1.3901.00
92_D146_Q1.3861.00
178_E181_Q1.3811.00
100_I121_V1.3771.00
346_L399_A1.3771.00
379_K383_S1.3771.00
53_L85_H1.3761.00
245_T466_N1.3641.00
280_V286_V1.3641.00
180_M206_I1.3481.00
24_N218_G1.3471.00
376_E379_K1.3361.00
227_T231_E1.3361.00
142_S145_M1.3281.00
122_Q134_P1.3271.00
15_P18_V1.3211.00
282_A421_D1.3141.00
11_R58_R1.3141.00
231_E234_S1.3101.00
284_E470_R1.3101.00
96_V132_I1.3091.00
95_T98_E1.3021.00
13_A57_E1.3001.00
264_V310_D1.2901.00
252_P281_K1.2891.00
133_H136_A1.2861.00
273_V302_A1.2841.00
9_N58_R1.2831.00
264_V302_A1.2831.00
419_N450_A1.2781.00
263_T311_S1.2681.00
264_V273_V1.2591.00
366_Y380_K1.2591.00
124_I156_Y1.2581.00
350_F354_E1.2521.00
111_R371_V1.2501.00
185_N188_K1.2501.00
383_S386_T1.2361.00
260_D279_S1.2361.00
51_F83_M1.2351.00
470_R482_S1.2281.00
210_V224_V1.2261.00
278_L472_A1.2151.00
393_P397_T1.2131.00
201_K229_Q1.2121.00
278_L288_I1.2121.00
443_K446_E1.2071.00
27_V161_I1.2041.00
325_T328_K1.1991.00
378_Y396_Y1.1941.00
222_L226_D1.1891.00
204_M229_Q1.1851.00
261_G277_S1.1821.00
398_H401_A1.1811.00
288_I472_A1.1751.00
218_G221_T1.1501.00
282_A453_A1.1501.00
437_I460_E1.1421.00
209_R223_D1.1351.00
126_D131_Q1.1341.00
271_E274_R1.1321.00
15_P19_A1.1271.00
204_M228_N1.1271.00
352_I410_A1.1251.00
458_S464_I1.1211.00
29_K160_D1.1181.00
28_K31_E1.1171.00
183_M193_I1.1141.00
183_M206_I1.1131.00
184_K188_K1.1031.00
327_R331_E1.1031.00
70_I75_K1.0971.00
222_L231_E1.0951.00
264_V272_T1.0951.00
411_I414_R1.0951.00
394_D397_T1.0931.00
297_S339_Q1.0921.00
464_I468_S1.0921.00
212_V235_L1.0911.00
337_I425_A1.0881.00
356_I381_A1.0881.00
380_K417_D1.0871.00
108_K113_D1.0861.00
389_D439_F1.0821.00
286_V482_S1.0821.00
164_F193_I1.0811.00
36_L200_L1.0761.00
97_A134_P1.0681.00
275_D478_R1.0671.00
248_R449_D1.0651.00
181_Q185_N1.0631.00
29_K64_R1.0631.00
26_Q209_R1.0631.00
56_P69_H1.0611.00
419_N452_K1.0561.00
464_I471_I1.0541.00
20_N49_V1.0531.00
211_T221_T1.0481.00
353_G378_Y1.0471.00
33_H221_T1.0431.00
23_I213_I1.0431.00
128_Y152_L1.0411.00
273_V276_L1.0401.00
353_G377_M1.0391.00
338_P344_H1.0391.00
261_G311_S1.0331.00
353_G395_E1.0311.00
4_V161_I1.0281.00
100_I156_Y1.0251.00
351_P354_E1.0241.00
48_N54_Y1.0231.00
75_K79_L1.0231.00
262_I302_A1.0161.00
106_P117_A1.0131.00
278_L286_V1.0121.00
475_F480_I1.0061.00
118_G122_Q1.0061.00
259_I303_V1.0041.00
347_V350_F1.0021.00
159_A162_L1.0001.00
27_V192_S0.9981.00
204_M224_V0.9911.00
5_I63_V0.9891.00
11_R21_D0.9881.00
286_V455_L0.9881.00
68_V79_L0.9871.00
329_I334_I0.9861.00
260_D277_S0.9861.00
254_A317_N0.9851.00
445_I449_D0.9851.00
427_Q459_F0.9851.00
384_L390_V0.9821.00
442_K446_E0.9821.00
323_N328_K0.9781.00
96_V141_I0.9781.00
87_H342_H0.9731.00
123_D126_D0.9691.00
444_L456_L0.9691.00
28_K209_R0.9681.00
356_I410_A0.9661.00
256_V282_A0.9651.00
87_H430_R0.9651.00
288_I455_L0.9631.00
8_L24_N0.9621.00
380_K383_S0.9591.00
115_K119_Q0.9571.00
282_A317_N0.9491.00
106_P120_E0.9471.00
424_I457_L0.9471.00
82_G154_T0.9431.00
134_P137_K0.9411.00
447_Q452_K0.9401.00
288_I299_L0.9361.00
423_L452_K0.9341.00
262_I309_T0.9341.00
55_Q69_H0.9311.00
162_L193_I0.9311.00
342_H348_L0.9311.00
4_V64_R0.9251.00
4_V63_V0.9211.00
335_G415_E0.9191.00
381_A395_E0.9181.00
304_T424_I0.9171.00
337_I411_I0.9161.00
296_Q426_A0.9141.00
393_P401_A0.9131.00
330_T415_E0.9111.00
125_S152_L0.9111.00
323_N334_I0.9111.00
177_K181_Q0.9101.00
158_G189_E0.9081.00
246_E466_N0.9071.00
119_Q122_Q0.9041.00
183_M195_L0.9041.00
203_I224_V0.8981.00
5_I50_L0.8951.00
74_N78_D0.8951.00
333_G421_D0.8951.00
367_S370_G0.8941.00
304_T415_E0.8931.00
352_I402_L0.8931.00
96_V136_A0.8891.00
12_K57_E0.8891.00
209_R221_T0.8881.00
82_G159_A0.8851.00
338_P411_I0.8841.00
323_N329_I0.8801.00
22_N58_R0.8771.00
444_L467_L0.8761.00
335_G423_L0.8741.00
359_Q415_E0.8741.00
97_A117_A0.8741.00
26_Q221_T0.8731.00
77_N157_R0.8731.00
61_I70_I0.8701.00
101_I120_E0.8661.00
259_I314_I0.8651.00
101_I117_A0.8641.00
353_G381_A0.8621.00
334_I422_L0.8601.00
273_V299_L0.8591.00
297_S426_A0.8541.00
63_V68_V0.8541.00
120_E158_G0.8531.00
382_R385_I0.8521.00
438_E442_K0.8481.00
210_V222_L0.8481.00
154_T162_L0.8461.00
33_H209_R0.8461.00
357_L361_Y0.8451.00
253_G282_A0.8411.00
210_V232_L0.8401.00
444_L454_V0.8391.00
424_I455_L0.8381.00
50_L61_I0.8351.00
384_L417_D0.8351.00
348_L399_A0.8301.00
49_V56_P0.8271.00
164_F183_M0.8261.00
468_S485_P0.8261.00
104_K156_Y0.8221.00
264_V270_I0.8221.00
122_Q135_E0.8181.00
51_F85_H0.8171.00
270_I310_D0.8171.00
276_L299_L0.8121.00
49_V54_Y0.8111.00
104_K157_R0.8081.00
130_L148_R0.8071.00
107_K113_D0.8041.00
32_I184_K0.7991.00
375_G379_K0.7951.00
46_L213_I0.7951.00
378_Y395_E0.7941.00
231_E235_L0.7941.00
360_S417_D0.7930.99
32_I187_V0.7930.99
342_H407_Q0.7920.99
22_N217_K0.7890.99
354_E377_M0.7880.99
124_I152_L0.7870.99
335_G420_P0.7840.99
278_L299_L0.7830.99
23_I218_G0.7820.99
129_G185_N0.7800.99
7_M25_L0.7780.99
107_K110_G0.7770.99
479_I482_S0.7760.99
296_Q339_Q0.7720.99
368_A373_H0.7720.99
118_G134_P0.7700.99
8_L26_Q0.7690.99
27_V194_I0.7670.99
106_P112_I0.7660.99
285_I445_I0.7650.99
279_S482_S0.7630.99
82_G157_R0.7630.99
412_I440_V0.7620.99
392_T395_E0.7610.99
276_L479_I0.7610.99
13_A19_A0.7590.99
90_L146_Q0.7590.99
138_A149_A0.7580.99
345_G411_I0.7570.99
264_V307_R0.7550.99
326_P329_I0.7530.99
70_I76_A0.7520.99
356_I413_G0.7520.99
93_T137_K0.7500.99
465_M471_I0.7470.99
89_M150_E0.7470.99
121_V134_P0.7440.99
257_L280_V0.7400.99
346_L407_Q0.7390.99
378_Y382_R0.7370.99
328_K331_E0.7370.99
348_L400_R0.7350.99
108_K111_R0.7350.99
376_E380_K0.7340.99
292_D459_F0.7330.99
28_K223_D0.7320.99
256_V422_L0.7310.99
122_Q133_H0.7310.99
170_V292_D0.7300.99
95_T137_K0.7300.99
365_P419_N0.7290.99
367_S371_V0.7270.99
328_K332_S0.7260.99
351_P396_Y0.7250.99
98_E105_E0.7240.99
15_P393_P0.7230.99
204_M227_T0.7230.99
88_F143_V0.7230.99
392_T402_L0.7210.99
48_N53_L0.7210.99
97_A106_P0.7210.99
162_L191_K0.7200.99
197_T203_I0.7150.99
387_E443_K0.7140.99
127_R158_G0.7140.99
288_I300_I0.7130.99
441_H466_N0.7090.99
316_L334_I0.7080.99
323_N332_S0.7060.99
101_I121_V0.7040.99
7_M61_I0.7040.99
263_T273_V0.6990.99
232_L236_M0.6980.99
234_S238_G0.6980.99
141_I149_A0.6980.99
264_V309_T0.6940.99
247_K445_I0.6930.99
48_N85_H0.6920.99
150_E153_K0.6910.99
125_S134_P0.6910.99
299_L303_V0.6910.99
386_T389_D0.6880.99
122_Q131_Q0.6880.99
34_A224_V0.6860.99
201_K205_E0.6840.99
465_M485_P0.6810.98
288_I474_I0.6810.98
335_G418_R0.6800.98
321_I332_S0.6790.98
258_A318_G0.6790.98
106_P116_R0.6780.98
124_I127_R0.6750.98
309_T322_Q0.6730.98
120_E124_I0.6720.98
74_N77_N0.6700.98
96_V149_A0.6690.98
254_A281_K0.6680.98
248_R469_D0.6680.98
114_R118_G0.6670.98
105_E117_A0.6630.98
264_V271_E0.6630.98
100_I149_A0.6630.98
85_H89_M0.6630.98
25_L33_H0.6620.98
351_P398_H0.6610.98
420_P452_K0.6590.98
257_L422_L0.6590.98
179_L183_M0.6580.98
195_L206_I0.6560.98
182_I185_N0.6560.98
187_V193_I0.6560.98
20_N213_I0.6560.98
35_L194_I0.6560.98
28_K192_S0.6560.98
304_T336_H0.6530.98
97_A121_V0.6530.98
302_A307_R0.6520.98
36_L203_I0.6510.98
90_L149_A0.6500.98
223_D227_T0.6500.98
256_V280_V0.6500.98
412_I432_L0.6480.98
129_G181_Q0.6470.98
352_I385_I0.6470.98
47_M163_I0.6470.98
70_I81_I0.6470.98
447_Q454_V0.6460.98
250_A253_G0.6450.98
262_I314_I0.6440.98
252_P255_E0.6430.98
390_V413_G0.6430.98
420_P423_L0.6420.98
422_L455_L0.6420.98
359_Q418_R0.6410.98
97_A135_E0.6390.98
357_L380_K0.6390.98
232_L238_G0.6390.98
124_I128_Y0.6380.98
184_K206_I0.6380.98
230_D234_S0.6370.98
263_T274_R0.6370.98
447_Q450_A0.6360.98
136_A145_M0.6340.98
247_K469_D0.6340.98
384_L387_E0.6300.97
352_I399_A0.6300.97
379_K382_R0.6290.97
141_I145_M0.6280.97
255_E260_D0.6280.97
304_T330_T0.6270.97
314_I323_N0.6260.97
88_F150_E0.6250.97
68_V71_N0.6250.97
118_G121_V0.6240.97
5_I161_I0.6230.97
259_I309_T0.6230.97
17_I393_P0.6220.97
257_L261_G0.6210.97
316_L332_S0.6210.97
445_I469_D0.6200.97
213_I218_G0.6190.97
300_I426_A0.6170.97
360_S365_P0.6160.97
174_H177_K0.6160.97
119_Q126_D0.6140.97
173_P177_K0.6130.97
7_M163_I0.6120.97
235_L238_G0.6110.97
92_D139_A0.6100.97
107_K111_R0.6100.97
100_I124_I0.6090.97
253_G281_K0.6080.97
237_V241_V0.6080.97
360_S418_R0.6050.97
215_K391_R0.6050.97
13_A21_D0.6040.97
214_R219_I0.6040.97
212_V232_L0.6030.97
380_K384_L0.6030.97
88_F169_A0.6030.97
397_T400_R0.6010.97
300_I336_H0.6010.97
298_E307_R0.6000.97
6_E65_G0.5990.97
96_V134_P0.5990.97
125_S130_L0.5990.97
106_P113_D0.5960.96
456_L468_S0.5950.96
381_A385_I0.5950.96
4_V29_K0.5950.96
200_L203_I0.5940.96
300_I455_L0.5940.96
250_A449_D0.5900.96
448_R454_V0.5900.96
63_V79_L0.5890.96
287_G458_S0.5890.96
172_T176_I0.5880.96
374_K377_M0.5880.96
471_I485_P0.5870.96
143_V430_R0.5870.96
181_Q188_K0.5860.96
384_L388_Y0.5850.96
224_V227_T0.5830.96
304_T326_P0.5830.96
33_H194_I0.5830.96
412_I416_I0.5830.96
473_V483_V0.5830.96
73_P76_A0.5820.96
15_P21_D0.5820.96
391_R397_T0.5820.96
338_P345_G0.5800.96
180_M187_V0.5800.96
337_I412_I0.5800.96
88_F170_V0.5790.96
25_L35_L0.5790.96
27_V33_H0.5780.96
19_A393_P0.5780.96
388_Y416_I0.5770.96
292_D296_Q0.5770.96
136_A141_I0.5760.96
176_I199_K0.5750.96
286_V453_A0.5750.96
131_Q145_M0.5750.96
399_A411_I0.5750.96
388_Y443_K0.5740.96
249_A252_P0.5730.96
143_V169_A0.5730.96
129_G178_E0.5720.96
296_Q459_F0.5710.96
12_K54_Y0.5710.96
263_T271_E0.5710.96
186_L191_K0.5710.96
161_I192_S0.5700.95
412_I425_A0.5700.95
239_R245_T0.5680.95
289_A461_L0.5680.95
25_L194_I0.5680.95
49_V57_E0.5680.95
348_L414_R0.5680.95
262_I273_V0.5680.95
36_L210_V0.5670.95
195_L203_I0.5670.95
168_T176_I0.5650.95
38_E239_R0.5650.95
172_T175_E0.5650.95
230_D240_E0.5640.95
278_L455_L0.5640.95
15_P397_T0.5630.95
38_E236_M0.5620.95
261_G313_T0.5610.95
279_S282_A0.5600.95
187_V206_I0.5600.95
276_L474_I0.5590.95
349_D398_H0.5570.95
352_I401_A0.5570.95
399_A414_R0.5570.95
243_F246_E0.5560.95
169_A430_R0.5560.95
255_E315_T0.5560.95
367_S373_H0.5550.95
388_Y413_G0.5530.95
61_I68_V0.5530.95
181_Q184_K0.5530.95
441_H445_I0.5510.95
285_I467_L0.5510.95
229_Q232_L0.5510.95
181_Q205_E0.5510.95
32_I208_D0.5510.95
281_K284_E0.5510.95
255_E259_I0.5510.95
330_T359_Q0.5510.95
238_G241_V0.5510.95
288_I479_I0.5500.95
32_I206_I0.5490.94
195_L199_K0.5490.94
72_S76_A0.5490.94
128_Y148_R0.5490.94
155_L186_L0.5490.94
301_E306_L0.5470.94
270_I274_R0.5460.94
207_C210_V0.5460.94
473_V480_I0.5450.94
439_F442_K0.5440.94
16_G19_A0.5430.94
35_L47_M0.5420.94
316_L421_D0.5420.94
176_I206_I0.5420.94
352_I397_T0.5420.94
360_S380_K0.5410.94
356_I385_I0.5400.94
31_E192_S0.5400.94
441_H467_L0.5390.94
302_A308_K0.5390.94
200_L232_L0.5380.94
239_R462_E0.5380.94
99_N138_A0.5380.94
47_M50_L0.5370.94
172_T292_D0.5370.94
63_V81_I0.5370.94
7_M24_N0.5350.94
289_A458_S0.5350.94
143_V146_Q0.5340.94
420_P425_A0.5340.94
356_I384_L0.5330.94
220_K227_T0.5320.94
331_E362_Y0.5310.93
304_T329_I0.5300.93
104_K120_E0.5270.93
314_I321_I0.5270.93
365_P368_A0.5270.93
101_I124_I0.5260.93
474_I479_I0.5260.93
144_G292_D0.5230.93
87_H150_E0.5220.93
25_L192_S0.5220.93
324_L329_I0.5220.93
354_E357_L0.5210.93
7_M49_V0.5190.93
236_M239_R0.5190.93
132_I152_L0.5180.93
18_V22_N0.5170.93
482_S485_P0.5170.93
345_G414_R0.5160.92
56_P67_K0.5150.92
163_I194_I0.5150.92
132_I136_A0.5150.92
114_R117_A0.5140.92
20_N23_I0.5140.92
27_V211_T0.5140.92
317_N320_Q0.5130.92
112_I117_A0.5120.92
395_E398_H0.5120.92
262_I277_S0.5120.92
336_H344_H0.5110.92
117_A120_E0.5090.92
173_P201_K0.5090.92
356_I417_D0.5090.92
271_E310_D0.5090.92
439_F443_K0.5070.92
438_E466_N0.5060.92
91_V94_F0.5050.92
282_A286_V0.5040.92
258_A313_T0.5040.92
13_A16_G0.5030.91
33_H192_S0.5030.91
121_V125_S0.5030.91
143_V170_V0.5010.91
392_T401_A0.5010.91
472_A481_A0.5010.91
44_S48_N0.5000.91
Legend: The i and j are positions as given in the UniProt sequence. Only predictions with Prob ≥ 0.7 are listed. For all 3L/2 predictions, click on one of the links below.

Sco = ((raw_sco - min_sco)/(avg_sco - min_sco))/2 + 0.5
Prob = P(contact | Sco, seq80/√len)
seq80 = effective number of sequences, after reducing redundancy to 80% identity

HHsearch Results:
Legend: GREMLIN results (Prob ≥ 0.7) overlaid on top PDB hits. The grey/red filled circles underneath are pdb residue contacts (min distance < 5 Angstroms). The shade of the circles is based on the top 10 (or top 20 if e-value < 1E-20) HHsearch results which uses the overall probability and per-site alignment prob. Inter oligomeric contacts in the pdb are in shades of red.
PDB Chains Cov % HHΔ  
4q9hA 1 0.951 100 0.403 Contact Map
4finA 2 0.9039 100 0.414 Contact Map
4f4cA 1 0.9333 100 0.429 Contact Map
1yqtA 1 0.9 100 0.45 Contact Map
3ux8A 2 0.8549 100 0.458 Contact Map
3bk7A 1 0.9176 100 0.46 Contact Map
3ozxA 1 0.8902 100 0.461 Contact Map
2iw3A 2 0.8882 100 0.469 Contact Map
3j16B 1 0.9196 100 0.475 Contact Map
2vf7A 2 0.9392 100 0.546 Contact Map

Sequence Conservation (Image Generated using WebLogo v3)
Legend: Color is used to show Hydrophobicity (Blue: Hydrophilic, Green: Neutral and Black: Hydrophobic). Amount of gaps is represented by width.

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